From the name, one of the points of the image is the open space. Visual narrative was supposed to go something like this:
That’s weird, there’s only stuff on the sides. Hey, birds! Boy that one’s really hanging out there. Oh, he’s looking across the space at a nest. Hey, more birds! Holy frack, this is a shocking surprise! Ah, the poor little fellows would probably like to get together – see how they’re looking at each other. Awww, poor birds. Oh wait. They’re birds. They can fly. Duh. OMG, another bird! Will the jubilation never end!?
Okay, maybe that’s pushing it as far as visual narrative goes, but you get the point.
As for the title using “whole”, well, there are several holes in the image, but at the same time, the bird family is whole, because gaps in space are nothing to them.