If AE doesn’t recognize the setting accurately, you can set it up yourself.
You can change the properties of the image source at interpret footage → main.
Take Control of Your After Effects Compositions (schoolofmotion.com)
And you can do it with composition setting.
(If the sauce is not accurate, there may be some problems.)
※ You can make the settings the same as the blender.
Yeah I’m really not sure what’s happening here… 
I have screenshot of the plane as it appears in the animation in the camera window…
And you can see the plane box from the Blender to AE is completely out of sync with the animation video… but have the same frame rate. The issue seems to be the size and ratio…
I got tagged … might be able to assist slightly.
Thanks for providing lots of screenshots, those really help in quickly getting a sense of what you’re dealing with.
I tried the AE export addon that comes with Blender many months ago; I can’t recall specifically what I was testing (I think it was something similar, doing billboard replacement) but I think it didn’t go well. I can’t remember the particulars exactly of what wasn’t working to well, but nonetheless your issue here doesn’t strike me as “it all works fine, the problem is you.”
The solution I’m using for AE isn’t free, but I’ve been rather pleased with it:
It does a good job of exporting various things - animated camera track, nulls, light information, etc. There’s a step or two that I’ve found has to be done perfectly, otherwise it gets the coordinates wrong.
Example: you cannot export nulls, then change your camera in blender, and export it by itself - it won’t sync. You need to export EVERYTHING in one go. So if you change anything about what your’re sending to AE, you need to export ALL of it again. This only takes mere seconds for export, but obviously anything you’ve done with the result in AE will require a rework - so, check all your tracking, before you start getting artsy with things.
Once i figured that out, everything was fairly painless. The plugin does come with a trial version, so I’d recommend you download it and play for a bit - see if it helps solve your issue. 
Ah ok I’ll give it a look and see if it helps my situation I’ll get back to you
Brilliant YES that fixes the problem 100% going to purchase the plugin now Thank you!
The location where Null is created in AE is based on blender’s Origins.
We do not use the generated Null, but rather use the coordinate value that the Null has.
Add a video to the project, copy Null’s coordinates and paste it into the video.
This will place the video in the Null position. 
※If the Null is a moving animation, the video must be connected to the Null so that it moves together.
