Skinning Not Working as Expected

I’m modeling and animating a tentacled creature, and ran into a problem with skinning the armature. Here’s a picture:

As you can see, the armature bones representing the tentacle move very much as expected with the IK constraint. However, the associated tentacle in the mesh gets all twisted. It’s like the movement/rotation of the bone is having too much of an effect on the mesh.

Does anyone know why?

Can’t really tell anything from a picture, upload the .blend file…


Here’s the blend file:


OK Thanks. The extra modifier must have been there because I detached a previous armature.

Also, that box showed up after I hit some key by accident, and I didn’t know what it was. Thanks for that.